This species have the following characteristics: pyloric caeca usually 7-8 in 1 or 2 whorls, typically 6+2; usually 3 rudimentary gill rakers on the upper arm of the first arch, 4-5 developed rakers on lower arm, usually 2-3 rudimentary rakers anteriorly on the lower arm; total gill rakers 9-13; dorsal-fin origin between neural spines 3-4; head and body uniformly dusky; pectoral fins clear; dorsal and anal fins clear with a dark edge, most developed in the anal fin; posterior part of the roof of the mouth, upper pharyngeal region, and gill bars are blackish; esophagus, caeca, and anterior part of the intestine are blackish (Ref. 91765).