Body oval; upper eye less than its own diameter from dorsal profile of head; anterior nostril on blind side not enlarged; anterior nostril on eyed side with a backward-pointing tube hardly reaching to front border of lower eye; dorsal fin beginning near vertical through front border of upper eye; pectoral fin on eyed side with 6-8 fin rays, on blind side shorter; caudal fin separate from last dorsal and anal fin rays; lateral line supra-temporal branch forming an angular S-shape, the ascending part running upward and backward (Ref. 4710).