Distinguished from all other species of the genus by the following combination of characters:

scales on the longitudinal series 21-22; a dark spot on dorsal fin above trunk bar 5, and usually

above trunk bar 4; head depth 90.2-96.3 % mm HL; cycloid scales on head sides; dorsal-fin rays 9-10; anal-fin rays 8-9; total vertebrae 24; scales on the upper lateral line 14-15; scales on the lower lateral line 6-8; absence of a dark dot at the edge of each scale on the midlateral portion of the flank; and a wide ectopterygoid (Ref. 80972).