In breeding males dorsal fin dusky to black, not yellow or orange; band on caudal fin narrow (narrower that eye-diameter) and often diffuse or absent; first dorsal fin ray thickened and spikelike; ocellus usually well developed in dorsal fin of females, no anal fin ocellus; breast fully scaled; modally 10 dorsal and anal fin rays; 7 pelvic fin rays, 25-28 (mode 26) scales in lateral line and modally 26 vertebrae (Ref. 43452).