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|Sound produced||yes, active sound production|
|Type of sound produced||grunts|
|Sound production organ||swim bladder|
|Sound mechanism||vibration by contraction of the drumming muscles associcated with the swimbladder|
|Behavioural context||courtship sound presumably by males during spawning period|
|Reference||Finstad, J.L. and J.T. Nordeide, 2001|
|Remark||recording was carried out in land-based fiberglass tanks during spawning period|