| Main Ref. | Sokolov, L.I. and L.S. Berdicheskii, 1989 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | one clear seasonal peak per year |
| Spawning aggregation | Yes. Ref. SCRFA, Science and Conservation of Fish Aggregations, 2018 |
| Batch spawner | Ref. Gerstmeier, R. and T. Romig, 1998 |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Spawn in large aggregations in fast flowing water on gravel bottom or submerged vegetation. Semi-anadromous individuals begin spawning migration in November in Kura (Ref. 59043). Sticky eggs attach themselves to stones or plants (Ref. 9953). |
| Search for more references on reproduction | Scirus |