| Main Ref. | Miller, P.J., 1986 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | |
| Spawning aggregation | Ref. |
| Batch spawner | Yes. Ref. Kottelat, M. and J. Freyhof, 2007 |
| Reproductive guild |
guarders nesters |
| Parental Care | paternal |
| Description of life cycle and mating behavior | Males defend eggs in cavities under stones, plant material or molluscs shells (Ref. 59043). Females spawn several times during a season (Ref. 59043). |
| Search for more references on reproduction | Scirus |