| Main Ref. | Whitehead, P.J.P., 1985 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Upriver movement to spawning areas occur in late afternoon or at night (Ref. 4639). |
| Search for more references on reproduction | Scirus |