| Main Ref. | Koli, L., 1990 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | one clear seasonal peak per year |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Ascend a short distance up the rivers in shoals in late August to mid-October. Mature in the second year (lacustrine forms later) and the young ones descend in late summer (Ref. 4779). |
| Search for more references on reproduction | Scirus |