| Remarks |
The type locality in Malaysia is in a coastal heath forest. This habitat consists mainly of Melaleuca (Myrtaceae) stands on sandy and peaty substrate. The waters are tannin stained, slow flowing, and acidic (pH 4.5, as measured in February 2009). Syntopic species recorded from the type locality include: Boraras maculatus, Cyclocheilichthys apogon, Osteochilus spilurus, Parachela maculicauda, P. oxygastroides, Rasbora einthovenii, R. trilineata, Systomus johorensis, Trigonopoma gracile, T. pauciperforatum (Cyprinidae), Lepidocephalichthys furcatus, Pangio alcoides, P. semicincta (Cobitidae), Kryptopterus macrocephalus, Ompok leiacanthus, Wallago leerii (Siluridae), Hemibagrus nemurus, Pseudomystus leiacanthus (Bagridae), Clarias batrachus, C. meladerma (Clariidae), Parakysis verrucosa (Akysidae), Aplocheilus panchax (Aplocheilidae), Hemirhamphodon pogonognathus (Hemiramphidae), Monopterus albus (Synbranchidae), Nandus nebulosus (Nandidae), Pristolepis grooti (Pristolepididae), Belontia hasseltii, Betta imbellis, B. waseri, Luciocephalus pulcher, Parosphromenus paludicola, Trichopodus leerii, T. trichopterus, Trichopsis vittata (Osphronemidae), Channa bankanensis, C. lucius and C. striata (Channidae) (Kottelat et al., 1992; unpublished data). |