Adult: 11.1-54.0 cm TL; body proportions, as percentage of TL (based on 39 specimens measuring 11.1-31.3 cm TL): 8.0-17.0 prebranchial length, 8.3-12.1 branchial length, 57.8-77.1 trunk length and 13.8-19.0 tail length. Mature males with large gular pouch that extends from immediately posterior of oral disc to end of branchial region; trunk myomeres, 78-84; dentition: supraoral lamina, 2 triangular supraoral laminae each with three unicuspid teeth, one per apex; infraoral lamina, 9-13 unicuspid teeth of various size in immature individuals and divided into 3 tricuspid plates with, in each case, the median cusp largest in spawning individuals; 7 endolateral plates on either side each bearing 3-5 unicuspid teeth (in prespawning individuals, usually 4-5); endolateral formula, typically 3-3-4-4-4-4-3 or 4-4-4-5-5-4-3 or 5 endolaterals as individual unicuspid teeth on either side in spawning individuals; 1-2 rows of anterials; first row of anterials consisting of 4 tricuspid radial plates in recently transformed individuals, of 2 tricuspid flanked on either side by 1 quadricuspid radial plate in feeding phase individuals and of 6 unicuspid teeth in spawning individuals; 1 row of exolaterals on each side; 1 row of posterials; first row of posterials, 10-12 tricuspid (sometimes quadricuspid) plates in prespawning individuals and 12 unicuspid teeth in spawning individuals; transverse lingual lamina, w-shaped, with numerous unicuspid teeth in prespawning individuals and with only 2 greatly enlarged lateral unicuspid teeth in spawning individuals; longitudinal lingual laminae, each with 9-15 unicuspid teeth; presence of caudal fin pigmentation but extent of coverage not determined; caudal fin shape, spade-like (Ref. 89241).